diff --git a/toonz/sources/include/toonz/tcleanupper.h b/toonz/sources/include/toonz/tcleanupper.h
index d6a1859f..3a2ac95a 100644
--- a/toonz/sources/include/toonz/tcleanupper.h
+++ b/toonz/sources/include/toonz/tcleanupper.h
@@ -57,7 +57,7 @@ public:
 
 class DVAPI TCleanupper {
   CleanupParameters *m_parameters;
-  TPointD m_customDpi;
+  TPointD m_sourceDpi;
 
 private:
   TCleanupper()
@@ -101,8 +101,8 @@ time, to unlock a possibly useful memory block.
   TRasterImageP autocenterOnly(const TRasterImageP &image, bool isCameraTest,
                                bool &autocentered);
 
-  TPointD getCustomDpi() const { return m_customDpi; }
-  void setCustomDpi(const TPointD &dpi) { m_customDpi = dpi; }
+  TPointD getSourceDpi() const { return m_sourceDpi; }
+  void setSourceDpi(const TPointD &dpi) { m_sourceDpi = dpi; }
 
 private:
   // process phase
diff --git a/toonz/sources/tcleanupper/tcleanupper.cpp b/toonz/sources/tcleanupper/tcleanupper.cpp
index b7503872..d22a1aa9 100644
--- a/toonz/sources/tcleanupper/tcleanupper.cpp
+++ b/toonz/sources/tcleanupper/tcleanupper.cpp
@@ -439,10 +439,19 @@ static void cleanupLevel(TXshSimpleLevel *xl, std::set<TFrameId> fidsInXsheet,
       updater.update(fid, ri);
       continue;
     }
-    double dpix, dpiy;
-    original->getDpi(dpix, dpiy);
-    cl->setCustomDpi((dpix == 0 && dpiy == 0) ? xl->getProperties()->getDpi()
-                                              : TPointD());
+    // Obtain the source dpi. Changed it to be done once at the first frame of
+    // each level in order to avoid the following problem:
+    // If the original raster level has no dpi (such as TGA images), obtaining
+    // dpi in every frame causes dpi mismatch between the first frame and the
+    // following frames, since the value
+    // TXshSimpleLevel::m_properties->getDpi() will be changed to the
+    // dpi of cleanup camera (= TLV's dpi) after finishing the first frame.
+    if (firstImage) {
+      TPointD dpi;
+      original->getDpi(dpi.x, dpi.y);
+      if (dpi.x == 0 && dpi.y == 0) dpi = xl->getProperties()->getDpi();
+      cl->setSourceDpi(dpi);
+    }
 
     CleanupPreprocessedImage *cpi;
     {
diff --git a/toonz/sources/toonz/cleanuppopup.cpp b/toonz/sources/toonz/cleanuppopup.cpp
index 252443d2..88604aae 100644
--- a/toonz/sources/toonz/cleanuppopup.cpp
+++ b/toonz/sources/toonz/cleanuppopup.cpp
@@ -292,9 +292,9 @@ CleanupPopup::CleanupPopup()
     , m_params(new CleanupParameters)
     , m_updater(new LevelUpdater)
     , m_originalLevelPath()
-    , m_originalPalette(0) {
+    , m_originalPalette(0)
+    , m_firstLevelFrame(true) {
   setWindowTitle(tr("Cleanup"));
-
   // Progress Bar
   m_progressLabel = new QLabel(tr("Cleanup in progress"));
   m_progressBar   = new QProgressBar;
@@ -1217,10 +1217,20 @@ void CleanupPopup::cleanupFrame() {
       IconGenerator::instance()->invalidate(sl, fid);
     } else {
       // Perform main processing
-      double dpix, dpiy;
-      original->getDpi(dpix, dpiy);
-      cl->setCustomDpi((dpix == 0 && dpiy == 0) ? sl->getProperties()->getDpi()
-                                                : TPointD());
+
+      // Obtain the source dpi. Changed it to be done once at the first frame of
+      // each level in order to avoid the following problem:
+      // If the original raster level has no dpi (such as TGA images), obtaining
+      // dpi in every frame causes dpi mismatch between the first frame and the
+      // following frames, since the value
+      // TXshSimpleLevel::m_properties->getDpi() will be changed to the
+      // dpi of cleanup camera (= TLV's dpi) after finishing the first frame.
+      if (m_firstLevelFrame) {
+        TPointD dpi;
+        original->getDpi(dpi.x, dpi.y);
+        if (dpi.x == 0 && dpi.y == 0) dpi = sl->getProperties()->getDpi();
+        cl->setSourceDpi(dpi);
+      }
 
       CleanupPreprocessedImage *cpi;
       {
diff --git a/toonz/sources/toonz/cleanupsettingsmodel.cpp b/toonz/sources/toonz/cleanupsettingsmodel.cpp
index 559caea2..499570a3 100644
--- a/toonz/sources/toonz/cleanupsettingsmodel.cpp
+++ b/toonz/sources/toonz/cleanupsettingsmodel.cpp
@@ -420,10 +420,10 @@ void CleanupSettingsModel::processFrame(TXshSimpleLevel *sl, TFrameId fid) {
   bool doCameraTest = (m_cameraTestsCount > 0);
 
   // Retrieve new image dpi
-  double dpix, dpiy;
-  imageToCleanup->getDpi(dpix, dpiy);
-  cl->setCustomDpi((dpix == 0 && dpiy == 0) ? sl->getProperties()->getDpi()
-                                            : TPointD());
+  TPointD dpi;
+  imageToCleanup->getDpi(dpi.x, dpi.y);
+  if (dpi.x == 0 && dpi.y == 0) dpi = sl->getProperties()->getDpi();
+  cl->setSourceDpi(dpi);
 
   // Perform primary cleanup processing
   if (doProcessing) {
diff --git a/toonz/sources/toonz/xshcellviewer.cpp b/toonz/sources/toonz/xshcellviewer.cpp
index c64c40cb..11d818a1 100644
--- a/toonz/sources/toonz/xshcellviewer.cpp
+++ b/toonz/sources/toonz/xshcellviewer.cpp
@@ -1332,13 +1332,13 @@ void CellArea::drawLevelCell(QPainter &p, int row, int col, bool isReference) {
   if (yetToCleanupCell)  // ORIENTATION: what's this?
   {
     if (o->isVerticalTimeline())
-      p.fillRect(
-          rect.adjusted(rect.width() / 2, 0, 0, 0),
-          (isSelected) ? SelectedFullcolorColumnColor : FullcolorColumnColor);
+      p.fillRect(rect.adjusted(rect.width() / 2, 0, 0, 0),
+                 (isSelected) ? m_viewer->getSelectedFullcolorColumnColor()
+                              : m_viewer->getFullcolorColumnColor());
     else
-      p.fillRect(
-          rect.adjusted(0, rect.height() / 2, 0, 0),
-          (isSelected) ? SelectedFullcolorColumnColor : FullcolorColumnColor);
+      p.fillRect(rect.adjusted(0, rect.height() / 2, 0, 0),
+                 (isSelected) ? m_viewer->getSelectedFullcolorColumnColor()
+                              : m_viewer->getFullcolorColumnColor());
   }
 
   bool isLastRow = nextCell.isEmpty() ||
diff --git a/toonz/sources/toonzlib/tcleanupper.cpp b/toonz/sources/toonzlib/tcleanupper.cpp
index 76475488..ab10c53c 100644
--- a/toonz/sources/toonzlib/tcleanupper.cpp
+++ b/toonz/sources/toonzlib/tcleanupper.cpp
@@ -416,11 +416,11 @@ bool TCleanupper::getResampleValues(const TRasterImageP &image, TAffine &aff,
        saveBox.getLy() > 0 && saveBox.getLy() < rasterLy))
     raster_is_savebox = false;
 
-  image->getDpi(dpi.x, dpi.y);
-  if (dpi == TPointD()) {
-    dpi                         = getCustomDpi();
-    if (dpi == TPointD()) dpi.x = dpi.y = 65.0;  // using 65.0 as default DPI
-  } else if (!dpi.x)
+  // Use the same source dpi throughout the level
+  dpi = getSourceDpi();
+  if (dpi == TPointD())
+    dpi.x = dpi.y = 65.0;  // using 65.0 as default DPI //??????WHY
+  else if (!dpi.x)
     dpi.x = dpi.y;
   else if (!dpi.y)
     dpi.y = dpi.x;